Charles Vacca, 39, was teaching the girl how to use the powerful weapon when he was shot in the head by a stray bullet.
It is believed the gun recoiled after the unnamed youngster pulled the trigger.
The former soldier was airlifted to Las Vegas' University Medical Centre where he was pronounced dead.
A video, which was released by police, shows the former soldier telling the youngster how to aim the gun.
He can be heard on the clip saying his final words: “All right, full auto."
As reported by The Telegraph, the girl was at the Last Stop outdoor shooting range in White Hills, Arizona, with her parents as part of a family holiday.
Speaking to NBC News, Sam Scarmardo, the manager at Last Stop range, said "the establish[ed] practice at most shooting ranges is eight-years-old and up with parental supervision."
Tributes have now been paid to Mr Vacca, who was married with a young family.
Writing on Facebook, his best friend Robert Vera said: “He became a brother and a major part of my life through thick and thin. Rest In Peace brother.”
Other friends described him as a "great guy" who was "very professional".
The range is popular with tourists and advertises “Burgers and Bullets” day. This includes lunch and a choice of more than 20 automatic weapons to shoot with.
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